2017 Ph.D. in Systematics and Evolution. Department of Biological Sciences, The George Washington University (advisor).
Faculty Mentor: Dr. R. Alexander Pyron
Dissertation: “Ecological and Evolutionary Drivers of Biodiversity and Extinction Risks in Amphibians and Squamates”
2011 M.Sc. in Animal Biology. Department of Biological Sciences, Universidade Federal do Espírito Santo
2009 B.Sc. in Biogeography. Department of Biological Sciences, Universidade Federal do Espírito Santo
2008 Licentiate in Biological Sciences. Department of Biological Sciences, Universidade Federal do Espírito Santo.
1. de Sá RO*, Tonini JFR*, Cuddy T, van Huss H, Long A, Forlani MC, Peloso PLV, Zaher H, Haddad CFB (2019) Multiple connections between Amazonia and Atlantic Forest shaped the phylogenetic and morphological diversity in the genus Chiasmocleis Mehely, 1904 (Anura: Microhylidae: Gastrophryninae). Molecular Phylogenetics and Evolution 130, p. 198–210. (*co-corresponding authors)
2. Shibatta LS, Tonini JFR, Abrahão VP, Oliveira C, Malabarba LR, Shibatta OA (2018) Reappraisal of the systematics of Microglanis cottoides (Siluriformes, Pseudopimelodidae), a catfish from southern Brazil. PLOS ONE 13(7): e0199963.
3. Tonini JFR, Beard K, Ferreira RB, Jetz W, Pyron RA (2016) Fully-sampled phylogenies of squamates reveal evolutionary patterns in threat status. Biological Conservation v. 204, p. 23–31.
See more at https://sites.google.com/view/joao-tonini/.